All 25 suspects identified by a probe into the devastating August 4 Beirut explosion are now in the custody of Lebanese authorities, a judicial source told AFP on Tuesday.

Authorities had already detained 21 suspects over the portside blast, which killed more than 180 people, wounded at least 6,500 others and wreaked devastation across the capital.

On Tuesday, lead investigative judge Fadi Sawan issued arrest warrants against the final four suspects after interrogating them, the judicial source said.

They include the head of military intelligence at the port, a major with the State Security agency, and two members of the General Security agency, he added.

The two security agencies, along with army and Lebanon’s customs authority, are the main bodies responsible for overseeing security at the port.

Hundreds of tonnes of ammonium nitrate, a highly explosive fertiliser, had been stored in a warehouse for at least six years, it emerged after the explosion.

That sparked widespread outrage over alleged official negligence that many said was to blame for the blast.

The exact circumstances leading up to the explosion remain unclear.

Security sources have suggested welding work could have started a fire that triggered the blast, but some observers have rejected this as a theory pushed by the authorities.

Lebanon has rejected calls for an international investigation into the country’s worst ever peacetime disaster, but the Lebanese probe is being aided by foreign experts, including from the US Federal Bureau of Investigation.

France, which counted several of its citizens among the dead, has launched its own inquiry.