An overladen ship crammed with thousands of sheep sank on Sunday in Sudan’s Red Sea port of Suakin, drowning most animals on board but with all crew surviving.

The livestock vessel was exporting the animals from Sudan to Saudi Arabia when it sank after several thousand more animals were loaded on board than it was meant to carry.

“The ship, Badr 1, sank during the early hours of Sunday morning,” a senior Sudanese port official said, speaking on condition of anonymity. “It was carrying 15,800 sheep, which was beyond its load limits.”

The official said the ship was supposed to carry only 9,000 sheep.

Another official, who said all crew were rescued, raised concerns over the economic and environmental impact of the accident.

“The sunken ship will affect the port’s operation,” the official said. “It will also likely have an environmental impact due to the death of the large number of animals carried by the ship.”

Omar al-Khalifa, head of the national exporters’ association, said the ship took several hours to sink at the pier – a window that meant it “could have been rescued”.

The total value of the lost livestock was $4m, said Saleh Selim, the head of the association’s livestock division, confirming also that the sheep were loaded onto the vessel at Suakin port.

He said livestock owners recovered only around 700 sheep “but they were found very ill and we don’t expect them to live long”.

Selim called for an investigation into the incident.

Last month, a massive fire broke out in the cargo area of Suakin port, lasting hours and causing heavy damage. It was not clear what caused the blaze.

An investigation has been launched to determine the cause of the fire, but has yet to release its findings.

The historic port town of Suakin is no longer Sudan’s main foreign trade hub, a role that has been taken by Port Sudan, 60km (40 miles) away along the Red Sea coast.

There have been moves to redevelop Suakin port, but a 2017 deal with Turkey to restore historic buildings and expand the docks was suspended after the ouster of longtime President Omar al-Bashir.

Sudan remains gripped by a chronic economic crisis, which has deepened following last year’s military coup led by army chief Abdel Fattah al-Burhan.

The military takeover triggered punitive measures, including aid cuts by Western governments, which demanded the restoration of the transitional administration installed after Bashir was toppled.